Normal gastric tissue Helicobacter pylori infection is associated with epigenetic age acceleration, increased mitotic tick rate, tissue cell composition, and Natural Killer cell methylation alterations

Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with . The mechanisms by which infection with contributes to carcinogenesis are not well understood. Recent studies from subjects with and without gastric cancer have identified significant DNA methyla...

Full description

Saved in:
Bibliographic Details
Published inbioRxiv
Main Authors Vlasac, Irma M, Christensen, Brock C, Salas, Lucas A
Format Journal Article Paper
LanguageEnglish
Published United States Cold Spring Harbor Laboratory 29.06.2023
Edition1.1
Subjects
Online AccessGet full text
ISSN2692-8205
2692-8205
DOI10.1101/2023.06.28.546926

Cover

Abstract Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with . The mechanisms by which infection with contributes to carcinogenesis are not well understood. Recent studies from subjects with and without gastric cancer have identified significant DNA methylation alterations in normal gastric mucosa associated with infection and gastric cancer risk. Here we further investigated DNA methylation alterations in normal gastric mucosa in gastric cancer cases (n = 42) and control subjects (n = 42) with infection data. We assessed tissue cell type composition, DNA methylation alterations within cell populations, epigenetic aging, and repetitive element methylation. In normal gastric mucosa of both gastric cancer cases and control subjects, we observed increased epigenetic age acceleration associated with infection. We also observed an increased mitotic tick rate associated with infection in both gastric cancer cases and controls. Significant differences in immune cell populations associated with infection in normal tissue from cancer cases and controls were identified using DNA methylation cell type deconvolution. We also found natural killer cell-specific methylation alterations in normal mucosa from gastric cancer patients with infection. Our findings from normal gastric mucosa provide insight into underlying cellular composition and epigenetic aspects of associated gastric cancer etiology.
AbstractList Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with . The mechanisms by which infection with contributes to carcinogenesis are not well understood. Recent studies from subjects with and without gastric cancer have identified significant DNA methylation alterations in normal gastric mucosa associated with infection and gastric cancer risk. Here we further investigated DNA methylation alterations in normal gastric mucosa in gastric cancer cases (n = 42) and control subjects (n = 42) with infection data. We assessed tissue cell type composition, DNA methylation alterations within cell populations, epigenetic aging, and repetitive element methylation. In normal gastric mucosa of both gastric cancer cases and control subjects, we observed increased epigenetic age acceleration associated with infection. We also observed an increased mitotic tick rate associated with infection in both gastric cancer cases and controls. Significant differences in immune cell populations associated with infection in normal tissue from cancer cases and controls were identified using DNA methylation cell type deconvolution. We also found natural killer cell-specific methylation alterations in normal mucosa from gastric cancer patients with infection. Our findings from normal gastric mucosa provide insight into underlying cellular composition and epigenetic aspects of associated gastric cancer etiology.
Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with Helicobacter pylori. The mechanisms by which infection with H. pylori contributes to carcinogenesis are not well understood. Recent studies from subjects with and without gastric cancer have identified significant DNA methylation alterations in normal gastric mucosa associated with H. pylori infection and gastric cancer risk. Here we further investigated DNA methylation alterations in normal gastric mucosa in gastric cancer cases (n = 42) and control subjects (n = 42) with H. pylori infection data. We assessed tissue cell type composition, DNA methylation alterations within cell populations, epigenetic aging, and repetitive element methylation.BackgroundGastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with Helicobacter pylori. The mechanisms by which infection with H. pylori contributes to carcinogenesis are not well understood. Recent studies from subjects with and without gastric cancer have identified significant DNA methylation alterations in normal gastric mucosa associated with H. pylori infection and gastric cancer risk. Here we further investigated DNA methylation alterations in normal gastric mucosa in gastric cancer cases (n = 42) and control subjects (n = 42) with H. pylori infection data. We assessed tissue cell type composition, DNA methylation alterations within cell populations, epigenetic aging, and repetitive element methylation.In normal gastric mucosa of both gastric cancer cases and control subjects, we observed increased epigenetic age acceleration associated with H. pylori infection. We also observed an increased mitotic tick rate associated with H. pylori infection in both gastric cancer cases and controls. Significant differences in immune cell populations associated with H. pylori infection in normal tissue from cancer cases and controls were identified using DNA methylation cell type deconvolution. We also found natural killer cell-specific methylation alterations in normal mucosa from gastric cancer patients with H. pylori infection.ResultsIn normal gastric mucosa of both gastric cancer cases and control subjects, we observed increased epigenetic age acceleration associated with H. pylori infection. We also observed an increased mitotic tick rate associated with H. pylori infection in both gastric cancer cases and controls. Significant differences in immune cell populations associated with H. pylori infection in normal tissue from cancer cases and controls were identified using DNA methylation cell type deconvolution. We also found natural killer cell-specific methylation alterations in normal mucosa from gastric cancer patients with H. pylori infection.Our findings from normal gastric mucosa provide insight into underlying cellular composition and epigenetic aspects of H. pylori associated gastric cancer etiology.ConclusionsOur findings from normal gastric mucosa provide insight into underlying cellular composition and epigenetic aspects of H. pylori associated gastric cancer etiology.
Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with Helicobacter pylori. The mechanisms by which infection with H. pylori contributes to carcinogenesis are not well understood. Recent studies from subjects with and without gastric cancer have identified significant DNA methylation alterations in normal gastric mucosa associated with H. pylori infection and gastric cancer risk. Here we further investigated DNA methylation alterations in normal gastric mucosa in gastric cancer cases (n = 42) and control subjects (n = 42) with H. pylori infection data. We assessed tissue cell type composition, DNA methylation alterations within cell populations, epigenetic aging, and repetitive element methylation. In normal gastric mucosa of both gastric cancer cases and control subjects, we observed increased epigenetic age acceleration associated with H. pylori infection. We also observed an increased mitotic tick rate associated with H. pylori infection in both gastric cancer cases and controls. Significant differences in immune cell populations associated with H. pylori infection in normal tissue from cancer cases and controls were identified using DNA methylation cell type deconvolution. We also found natural killer cell-specific methylation alterations in normal mucosa from gastric cancer patients with H. pylori infection. Our findings from normal gastric mucosa provide insight into underlying cellular composition and epigenetic aspects of H. pylori associated gastric cancer etiology.
Author Vlasac, Irma M
Salas, Lucas A
Christensen, Brock C
Author_xml – sequence: 1
  givenname: Irma M
  orcidid: 0000-0002-1037-2698
  surname: Vlasac
  fullname: Vlasac, Irma M
– sequence: 2
  givenname: Brock C
  surname: Christensen
  fullname: Christensen, Brock C
– sequence: 3
  givenname: Lucas A
  surname: Salas
  fullname: Salas, Lucas A
BackLink https://www.ncbi.nlm.nih.gov/pubmed/37425894$$D View this record in MEDLINE/PubMed
BookMark eNpNkU1v1DAQhi1URD_oD-BS-cihu_gj9sZHVJUWUZULnKOJM9kanDi1nZb9if1X9Xa3wGVmJD_zvh69x-RgDCMS8oGzJeeMfxJMyCXTS1EvVaWN0G_IkSh9UQumDv6bD8lpSr8YY8JoLlfVO3JYqlC1qY7I022IA3i6hpSjszS7lGak1-idDS3YjJFOGx-io27s0WYXRuoShZSCdZCxo48u31Gc3BpHzEUC1kjBWvQYYYufl00bEVJhB5dDfrGxv2l5xvNXx8J7asMwheR2WzB29BbyHMv3vjlf5HbQgPlu41-kKfi8d0nvydsefMLTfT8hP79c_ri4Xtx8v_p68flm0fJKqQVX5fC-5tpyxbg1phetMdhaxRlIKSoted0bgFVvOjR9u-I91kzrrquUVK08IR93uq0L8Y97aKboBoibZhtIw3Qj6mYXyD90iuF-xpSbwaXtDTBimFMhpRFKKLVFz_bo3A7Y_RV9TUo-A32Mm7A
ContentType Journal Article
Paper
Copyright 2023, Posted by Cold Spring Harbor Laboratory
Copyright_xml – notice: 2023, Posted by Cold Spring Harbor Laboratory
DBID NPM
7X8
FX.
DOI 10.1101/2023.06.28.546926
DatabaseName PubMed
MEDLINE - Academic
bioRxiv
DatabaseTitle PubMed
MEDLINE - Academic
DatabaseTitleList PubMed
MEDLINE - Academic

Database_xml – sequence: 1
  dbid: FX.
  name: bioRxiv
  url: https://www.biorxiv.org/
  sourceTypes: Open Access Repository
– sequence: 2
  dbid: NPM
  name: PubMed
  url: https://proxy.k.utb.cz/login?url=http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=PubMed
  sourceTypes: Index Database
DeliveryMethod fulltext_linktorsrc
Discipline Biology
EISSN 2692-8205
Edition 1.1
ExternalDocumentID 2023.06.28.546926v1
37425894
Genre Preprint
GrantInformation_xml – fundername: NCI NIH HHS
  grantid: P30 CA023108
– fundername: NCI NIH HHS
  grantid: R01 CA253976
– fundername: NCI NIH HHS
  grantid: R01 CA216265
– fundername: NIGMS NIH HHS
  grantid: P20 GM104416
GroupedDBID NPM
7X8
8FE
8FH
AFKRA
ALMA_UNASSIGNED_HOLDINGS
BBNVY
BENPR
BHPHI
CCPQU
HCIFZ
LK8
M7P
NQS
PHGZM
PHGZT
PIMPY
PQGLB
PROAC
PUEGO
RHI
FX.
ID FETCH-LOGICAL-b1455-15589f816c1501c99f2b99ebc510a33246318f9aa7f9de9fb71fe8066dd4535b3
IEDL.DBID FX.
ISSN 2692-8205
IngestDate Tue Jan 07 18:49:12 EST 2025
Thu Sep 04 18:49:45 EDT 2025
Thu Apr 03 07:08:15 EDT 2025
IsDoiOpenAccess true
IsOpenAccess true
IsPeerReviewed false
IsScholarly false
Keywords epigenetic clocks
infection
cell proportions
DNA methylation
immune cells
Gastric cancer
epigenetics
age acceleration
Language English
License This pre-print is available under a Creative Commons License (Attribution-NonCommercial-NoDerivs 4.0 International), CC BY-NC-ND 4.0, as described at http://creativecommons.org/licenses/by-nc-nd/4.0
LinkModel DirectLink
MergedId FETCHMERGED-LOGICAL-b1455-15589f816c1501c99f2b99ebc510a33246318f9aa7f9de9fb71fe8066dd4535b3
Notes ObjectType-Article-2
SourceType-Scholarly Journals-1
ObjectType-Working Paper/Pre-Print-1
ObjectType-Feature-3
content type line 23
Competing Interest Statement: The authors have declared no competing interest.
ORCID 0000-0002-1037-2698
OpenAccessLink https://www.biorxiv.org/content/10.1101/2023.06.28.546926
PMID 37425894
PQID 2839252556
PQPubID 23479
PageCount 29
ParticipantIDs biorxiv_primary_2023_06_28_546926
proquest_miscellaneous_2839252556
pubmed_primary_37425894
PublicationCentury 2000
PublicationDate 2023-Jun-29
20230629
PublicationDateYYYYMMDD 2023-06-29
PublicationDate_xml – month: 06
  year: 2023
  text: 2023-Jun-29
  day: 29
PublicationDecade 2020
PublicationPlace United States
PublicationPlace_xml – name: United States
PublicationTitle bioRxiv
PublicationTitleAlternate bioRxiv
PublicationYear 2023
Publisher Cold Spring Harbor Laboratory
Publisher_xml – name: Cold Spring Harbor Laboratory
References Woo, Fernandez-Jimenez, Ghantous, Degli Esposti, Cuenin, Cahais (2023.06.28.546926v1.16) 2018; 143
Hannum, Guinney, Zhao, Zhang, Hughes, Sadda (2023.06.28.546926v1.20) 2013; 49
Breeze, Reynolds, van Dongen, Dunham, Lazar, Neph (2023.06.28.546926v1.26) 2019; 35
Ehrlich (2023.06.28.546926v1.8) 2002; 21
Kumar, Metz, Ellenberg, Kaplan, Goldberg (2023.06.28.546926v1.4) 2020; 158
Osaki, Bockerstett, Wong, Ford, Madison, DiPaolo (2023.06.28.546926v1.40) 2019; 247
Teschendorff, Marabita, Lechner, Bartlett, Tegner, Gomez-Cabrero (2023.06.28.546926v1.18) 2013; 29
Chung, Lim (2023.06.28.546926v1.5) 2014; 20
Tan, Yeoh (2023.06.28.546926v1.1) 2015; 149
Hong, Yang, Wang, Zhang, Wu, Chen (2023.06.28.546926v1.27) 2021
Kong, Rose, Cass, Williams, Darwish, Lianoglou (2023.06.28.546926v1.30) 2019; 10
Semper, Vieth, Gerhard, Mejías-Luque (2023.06.28.546926v1.42) 2019; 203
Berdasco, Esteller (2023.06.28.546926v1.7) 2010; 19
Ritchie, Phipson, Wu, Hu, Law, Shi (2023.06.28.546926v1.23) 2015; 43
Horvath (2023.06.28.546926v1.13) 2013; 14
Tomasetti, Vogelstein (2023.06.28.546926v1.15) 2015; 347
Robertson (2023.06.28.546926v1.10) 2005; 6
Xu, Niu, Li, Taylor (2023.06.28.546926v1.17) 2016; 44
Zhou, Laird, Shen (2023.06.28.546926v1.19) 2017; 45
Schoenborn, Wilson (2023.06.28.546926v1.41) 2007
Doglioni, Moschini, de Boni, Wotherspoon, Isaacson (2023.06.28.546926v1.32) 1992; 339
Qinyu, Long, Zhen-dong, Min-min, Wei-ze, Wei-ping (2023.06.28.546926v1.28) 2013; 587
Arlt, Schäfer, Kalthoff (2023.06.28.546926v1.38) 2012; 1
Zheng, Joyce, Liu, Zhang, Kibbe, Zhang (2023.06.28.546926v1.22) 2017; 45
Ushijima, Hattori (2023.06.28.546926v1.31) 2012; 18
Chiba, Marusawa, Seno, Watanabe (2023.06.28.546926v1.34) 2008; 23
DiLillo, Yanaba, Tedder (2023.06.28.546926v1.43) 2010; 184
Nishiyama, Nakanishi (2023.06.28.546926v1.9) 2021; 37
Bach, Long, Luu, Anh, Kwon, Lee (2023.06.28.546926v1.37) 2018; 19
Pappalardo, Barra (2023.06.28.546926v1.12) 2021; 14
Teschendorff (2023.06.28.546926v1.14) 2020; 12
Yamashita, Nanjo, Rehnberg, Iida, Takeshima, Ando (2023.06.28.546926v1.6) 2019; 11
Jan, Rather, Mushtaq, Malik, Besina, Baba (2023.06.28.546926v1.39) 2021
Slotkin, Martienssen (2023.06.28.546926v1.29) 2007; 8
Du, Wei (2023.06.28.546926v1.35) 2019; 9
Yun, Lundgren, Azem, Sjöling, Holmgren, Svennerholm (2023.06.28.546926v1.36) 2005; 73
Salas, Koestler, Butler, Hansen, Wiencke, Kelsey (2023.06.28.546926v1.25) 2018; 19
Alipour (2023.06.28.546926v1.2) 2021; 52
Levine, Lu, Quach, Chen, Assimes, Bandinelli (2023.06.28.546926v1.21) 2018; 10
Sung, Ferlay, Siegel, Laversanne, Soerjomataram, Jemal (2023.06.28.546926v1.3) 2021; 71
Zheng, Breeze, Beck, Teschendorff (2023.06.28.546926v1.24) 2018; 15
Du, Isaccson (2023.06.28.546926v1.33) 2002; 3
Niwa, Tsukamoto, Toyoda, Mori, Tanaka, Maekita (2023.06.28.546926v1.11) 2010; 70
References_xml – volume: 44
  start-page: e20
  issue: 3
  year: 2016
  ident: 2023.06.28.546926v1.17
  article-title: ENmix: a novel background correction method for Illumina HumanMethylation450 BeadChip
  publication-title: Nucleic Acids Res
– volume: 14
  start-page: 25
  issue: 1
  year: 2021
  ident: 2023.06.28.546926v1.12
  article-title: Losing DNA methylation at repetitive elements and breaking bad
  publication-title: Epigenetics & Chromatin
– volume: 43
  start-page: e47
  issue: 7
  year: 2015
  end-page: e47
  ident: 2023.06.28.546926v1.23
  article-title: . limma powers differential expression analyses for RNA-sequencing and microarray studies
  publication-title: Nucleic Acids Research
– volume: 21
  start-page: 5400
  issue: 35
  year: 2002
  end-page: 13
  ident: 2023.06.28.546926v1.8
  article-title: DNA methylation in cancer: too much, but also too little
  publication-title: Oncogene
– volume: 10
  start-page: 573
  issue: 4
  year: 2018
  end-page: 91
  ident: 2023.06.28.546926v1.21
  article-title: An epigenetic biomarker of aging for lifespan and healthspan
  publication-title: Aging (Albany NY)
– volume: 158
  start-page: 527
  issue: 3
  year: 2020
  end-page: 536
  ident: 2023.06.28.546926v1.4
  article-title: Risk Factors and Incidence of Gastric Cancer After Detection of Helicobacter pylori Infection: A Large Cohort Study
  publication-title: Gastroenterology
– volume: 70
  start-page: 1430
  issue: 4
  year: 2010
  end-page: 40
  ident: 2023.06.28.546926v1.11
  article-title: Inflammatory Processes Triggered by Helicobacter pylori Infection Cause Aberrant DNA Methylation in Gastric Epithelial Cells
  publication-title: Cancer Research
– volume: 23
  start-page: 1175
  issue: 8pt1
  year: 2008
  end-page: 81
  ident: 2023.06.28.546926v1.34
  article-title: Mechanism for gastric cancer development by Helicobacter pylori infection
  publication-title: Journal of Gastroenterology and Hepatology
– volume: 339
  start-page: 834
  issue: 8797
  year: 1992
  end-page: 5
  ident: 2023.06.28.546926v1.32
  article-title: High incidence of primary gastric lymphoma in northeastern Italy
  publication-title: The Lancet
– start-page: 41
  year: 2007
  end-page: 101
  ident: 2023.06.28.546926v1.41
  publication-title: Advances in Immunology [Internet]
– start-page: 12
  year: 2021
  ident: 2023.06.28.546926v1.27
  article-title: Epigenetic Age Acceleration of Stomach Adenocarcinoma Associated With Tumor Stemness Features, Immunoactivation, and Favorable Prognosis
  publication-title: Frontiers in Genetics [Internet]
– volume: 184
  start-page: 4006
  issue: 7
  year: 2010
  end-page: 16
  ident: 2023.06.28.546926v1.43
  article-title: B cells are required for optimal CD4+ and CD8+ T cell tumor immunity: therapeutic B cell depletion enhances B16 melanoma growth in mice
  publication-title: J Immunol
– volume: 18
  start-page: 923
  issue: 4
  year: 2012
  end-page: 9
  ident: 2023.06.28.546926v1.31
  article-title: Molecular Pathways: Involvement of Helicobacter pylori– Triggered Inflammation in the Formation of an Epigenetic Field Defect, and Its Usefulness as Cancer Risk and Exposure Markers
  publication-title: Clinical Cancer Research
– volume: 20
  start-page: 1667
  issue: 7
  year: 2014
  end-page: 80
  ident: 2023.06.28.546926v1.5
  article-title: Role of the tumor microenvironment in the pathogenesis of gastric carcinoma
  publication-title: World J Gastroenterol
– volume: 9
  start-page: 3095
  year: 2019
  ident: 2023.06.28.546926v1.35
  article-title: Therapeutic Potential of Natural Killer Cells in Gastric Cancer
  publication-title: Front Immunol
– volume: 6
  start-page: 597
  issue: 8
  year: 2005
  end-page: 610
  ident: 2023.06.28.546926v1.10
  article-title: DNA methylation and human disease
  publication-title: Nat Rev Genet
– volume: 8
  start-page: 272
  issue: 4
  year: 2007
  end-page: 85
  ident: 2023.06.28.546926v1.29
  article-title: Transposable elements and the epigenetic regulation of the genome
  publication-title: Nat Rev Genet
– volume: 19
  start-page: 698
  issue: 5
  year: 2010
  end-page: 711
  ident: 2023.06.28.546926v1.7
  article-title: Aberrant Epigenetic Landscape in Cancer: How Cellular Identity Goes Awry
  publication-title: Developmental Cell
– volume: 19
  start-page: 64
  year: 2018
  ident: 2023.06.28.546926v1.25
  article-title: An optimized library for reference-based deconvolution of whole-blood biospecimens assayed using the Illumina HumanMethylationEPIC BeadArray
  publication-title: Genome Biol
– volume: 45
  start-page: e22
  issue: 4
  year: 2017
  ident: 2023.06.28.546926v1.19
  article-title: Comprehensive characterization, annotation and innovative use of Infinium DNA methylation BeadChip probes
  publication-title: Nucleic Acids Res
– volume: 1
  start-page: e35
  issue: 11
  year: 2012
  end-page: e35
  ident: 2023.06.28.546926v1.38
  article-title: The ‘N-factors’ in pancreatic cancer: functional relevance of NF-κB, NFAT and Nrf2 in pancreatic cancer
  publication-title: Oncogenesis
– volume: 247
  start-page: 513
  issue: 4
  year: 2019
  end-page: 23
  ident: 2023.06.28.546926v1.40
  article-title: Interferon-γ directly induces gastric epithelial cell death and is required for progression to metaplasia
  publication-title: J Pathol
– volume: 49
  start-page: 359
  issue: 2
  year: 2013
  end-page: 67
  ident: 2023.06.28.546926v1.20
  article-title: Genome-wide methylation profiles reveal quantitative views of human aging rates
  publication-title: Mol Cell
– volume: 587
  start-page: 2105
  issue: 14
  year: 2013
  end-page: 11
  ident: 2023.06.28.546926v1.28
  article-title: FOXO6 promotes gastric cancer cell tumorigenicity via upregulation of C-myc
  publication-title: FEBS Letters
– volume: 52
  start-page: 23
  issue: 1
  year: 2021
  end-page: 30
  ident: 2023.06.28.546926v1.2
  article-title: Molecular Mechanism of Helicobacter pylori-Induced Gastric Cancer
  publication-title: J Gastrointest Canc
– start-page: 10
  year: 2021
  ident: 2023.06.28.546926v1.39
  article-title: Helicobacter pylori Subdues Cytokine Signaling to Alter Mucosal Inflammation via Hypermethylation of Suppressor of Cytokine Signaling 1 Gene During Gastric Carcinogenesis
  publication-title: Frontiers in Oncology [Internet]
– volume: 203
  start-page: 2183
  issue: 8
  year: 2019
  end-page: 93
  ident: 2023.06.28.546926v1.42
  article-title: Helicobacter pylori Exploits the NLRC4 Inflammasome to Dampen Host Defenses
  publication-title: The Journal of Immunology
– volume: 11
  start-page: 191
  issue: 1
  year: 2019
  ident: 2023.06.28.546926v1.6
  article-title: Distinct DNA methylation targets by aging and chronic inflammation: a pilot study using gastric mucosa infected with Helicobacter pylori
  publication-title: Clin Epigenet
– volume: 37
  start-page: 1012
  issue: 11
  year: 2021
  end-page: 27
  ident: 2023.06.28.546926v1.9
  article-title: Navigating the DNA methylation landscape of cancer
  publication-title: Trends in Genetics
– volume: 45
  start-page: 8697
  issue: 15
  year: 2017
  end-page: 711
  ident: 2023.06.28.546926v1.22
  article-title: Prediction of genome-wide DNA methylation in repetitive elements
  publication-title: Nucleic Acids Res
– volume: 19
  start-page: 3279
  issue: 10
  year: 2018
  ident: 2023.06.28.546926v1.37
  article-title: The Dominant Role of Forkhead Box Proteins in Cancer
  publication-title: International Journal of Molecular Sciences
– volume: 10
  start-page: 5228
  issue: 1
  year: 2019
  ident: 2023.06.28.546926v1.30
  article-title: Transposable element expression in tumors is associated with immune infiltration and increased antigenicity
  publication-title: Nat Commun
– volume: 149
  start-page: 1153
  issue: 5
  year: 2015
  end-page: 1162
  ident: 2023.06.28.546926v1.1
  article-title: Genetics and Molecular Pathogenesis of Gastric Adenocarcinoma
  publication-title: Gastroenterology
– volume: 73
  start-page: 1482
  issue: 3
  year: 2005
  end-page: 90
  ident: 2023.06.28.546926v1.36
  article-title: Natural Killer Cells and Helicobacter pylori Infection: Bacterial Antigens and Interleukin-12 Act Synergistically To Induce Gamma Interferon Production
  publication-title: Infection and Immunity
– volume: 14
  start-page: 3156
  issue: 10
  year: 2013
  ident: 2023.06.28.546926v1.13
  article-title: DNA methylation age of human tissues and cell types
  publication-title: Genome Biology
– volume: 347
  start-page: 78
  issue: 6217
  year: 2015
  end-page: 81
  ident: 2023.06.28.546926v1.15
  article-title: Variation in cancer risk among tissues can be explained by the number of stem cell divisions
  publication-title: Science
– volume: 35
  start-page: 4767
  issue: 22
  year: 2019
  end-page: 9
  ident: 2023.06.28.546926v1.26
  article-title: eFORGE v2.0: updated analysis of cell type-specific signal in epigenomic data
  publication-title: Bioinformatics
– volume: 71
  start-page: 209
  issue: 3
  year: 2021
  end-page: 49
  ident: 2023.06.28.546926v1.3
  article-title: Global Cancer Statistics 2020: GLOBOCAN Estimates of Incidence and Mortality Worldwide for 36 Cancers in 185 Countries
  publication-title: CA: A Cancer Journal for Clinicians
– volume: 3
  start-page: 97
  issue: 2
  year: 2002
  end-page: 104
  ident: 2023.06.28.546926v1.33
  article-title: Gastric MALT lymphoma: from aetiology to treatment
  publication-title: The Lancet Oncology
– volume: 12
  start-page: 56
  issue: 1
  year: 2020
  ident: 2023.06.28.546926v1.14
  article-title: A comparison of epigenetic mitotic-like clocks for cancer risk prediction
  publication-title: Genome Medicine
– volume: 15
  start-page: 1059
  issue: 12
  year: 2018
  end-page: 66
  ident: 2023.06.28.546926v1.24
  article-title: Identification of differentially methylated cell types in epigenome-wide association studies
  publication-title: Nat Methods
– volume: 143
  start-page: 597
  issue: 3
  year: 2018
  end-page: 609
  ident: 2023.06.28.546926v1.16
  article-title: Genome-wide profiling of normal gastric mucosa identifies Helicobacter pylori- and cancer-associated DNA methylome changes
  publication-title: Int J Cancer
– volume: 29
  start-page: 189
  issue: 2
  year: 2013
  end-page: 96
  ident: 2023.06.28.546926v1.18
  article-title: A beta-mixture quantile normalization method for correcting probe design bias in Illumina Infinium 450 k DNA methylation data
  publication-title: Bioinformatics
SSID ssj0002961374
Score 1.8395879
SecondaryResourceType preprint
Snippet Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with . The mechanisms by which infection with contributes to...
Gastric adenocarcinomas are a leading cause of global mortality, associated with chronic infection with Helicobacter pylori. The mechanisms by which infection...
SourceID biorxiv
proquest
pubmed
SourceType Open Access Repository
Aggregation Database
Index Database
SubjectTerms Cancer Biology
Title Normal gastric tissue Helicobacter pylori infection is associated with epigenetic age acceleration, increased mitotic tick rate, tissue cell composition, and Natural Killer cell methylation alterations
URI https://www.ncbi.nlm.nih.gov/pubmed/37425894
https://www.proquest.com/docview/2839252556
https://www.biorxiv.org/content/10.1101/2023.06.28.546926
hasFullText 1
inHoldings 1
isFullTextHit
isPrint
journalDatabaseRights – providerCode: PRVPQU
  databaseName: ProQuest Central
  customDbUrl: http://www.proquest.com/pqcentral?accountid=15518
  eissn: 2692-8205
  dateEnd: 99991231
  omitProxy: true
  ssIdentifier: ssj0002961374
  issn: 2692-8205
  databaseCode: BENPR
  dateStart: 20131107
  isFulltext: true
  titleUrlDefault: https://www.proquest.com/central
  providerName: ProQuest
link http://utb.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwjV1LSwMxEA7aInjzbX2UETx2Szf7zFVpKYpLEQu9LdkkK0XdlraK_kT_lTPJVjwoeFmWJdmQTJL5ZjL5hrFLQQwvXCo0U5X28NVeVk48FUdco37QvYjuDt9l8XAc3kyiyY9UXxRWWUxni_fpmz3Hp4Bt3H3d4u75ZKsHRLjJ026Elh2PN1kzQRVP0VyDSffbvcIF6qkkrM8xf62JiLdu6W90abXMYIc1R3JuFrtsw1R7bMulifzYZ58ZActneJSUZEPByg4WoMZAMRaWbhnmZHhPYR1bVcF0CbIeeqOB3K1g5kS9SbcWAXcRkEqhznEzoIM1CT8usewLLvKVbUY9ATFJdNYtkpcfKAi9jvTqgKw0ZNJyd8CtvVboClFi6g8XZgf2QN45Bg_YeNB_uB56dQoGryAGcw_RRirK1I8VAkdfCVHyQghTKFzKMkAwFuOeUAopk1JoI8oi8UuTIozROoyCqAgOWaOaVeaYAUnJF5LLEk0wY4wIA64TKRAkyDjtBS12UYsjnzuijZxElvfinKe5ExmWWQsqx2VAHZKVmb0usQgCvYj41FrsyEnw-zc4Dzh2Izz5RwunbJu-UTAYF2essVq8mnOEHauizZpX_Wx037YTDZ_Z6O4Ls1XYDA
linkProvider Cold Spring Harbor Laboratory Press
linkToHtml http://utb.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwjV3JTsMwELVYhODGTlkHiWNTNc7qM6IqW8QBpN4ix3ZQBaRVkyL4RP6KGTvlBBK3KBnHssf2vBmPnxm7EMTwwqVCN1VpDx_tYeXEU3HENdoH3Y_o7PB9Fg-fwptRNGoDbnWbVlmMJ7OP8bvdx6eEbVx93eTu--SrB0S4ydNehJ4dj3sUpl5mqwnaeRrhg1HvJ8aCb_wgCdvNzF-LI-xtq_sbYlpTM9hkqw9yamZbbMlU22zN3RX5ucO-MkKXr_As6aYNBY3tMUCzgbosLOcyTMn7HsMiwaqCcQ2y7X-jgWKuYKbEv0lHFwGXEpBKoeFxw6CLJQlE1ij7hjO9sdWoFyA6ie6iRgr1A2Wit-leXZCVhkxaAg-4tWcLnRDdTv3pcu3A7sq76OAuexpcPV4OvfYeBq8gGnMPIUcqytSPFaJHXwlR8kIIUyiczzJARBbjwlAKKZNSaCPKIvFLkyKW0TqMgqgI9thKNanMAQPSki8klyX6YcYYEQZcJ1IgUpBx2g867LxVRz51bBs5qSzvxzlPc6cylFkoKse5QA2SlZnMaxRBtBcRqVqH7TsN_vwGxwHHZoSH_6jhjK0PH-_v8rvr7PaIbdB3yg7j4pitNLO5OUEc0hSndrB9A14M2jQ
linkToPdf http://utb.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwjV1LT8MwDI6ACcSNN-NpJI7rtKbPnIEJGEw7gLRblSYpmoCu2gZiP5F_hZ10iANI3Co1TZQ6iT879mfGzgUxvHCp0ExV2sNHm6yceCqOuEb9oDsR5Q7f9-Prx_B2GA1_5MJQWGU-Gk8-Ru_2Hp8CtvH0dZu745OtHhDhJk_bEVp2PG6Tm7pd6WKZNRLU9WSAdYftbz8LF6iwkrC-0Py1C4S-9ZB_w0yrbrobrDGQlZlssiVTbrFVVy9yvs0--4QwX-BJUrUNBTP71wBVB8ozt7zLUJEFPoJFkFUJoynIWgZGA_ldwVTEwUnpi4DHCUilUPm4pdDCLwlITrHtK-72mR1GPQNRSrQWI5K7HygavQ75aoEsNfSlJfGAns0vdI2oQvXcxduBvZl3HsId9ti9eri49upaDF5OVOYewo5UFKkfK0SQvhKi4LkQJle4p2WAqCzGw6EQUiaF0EYUeeIXJkU8o3UYBVEe7LKVclyafQYkJV9ILgu0xYwxIgy4TqRAtCDjtBM02VktjqxyjBsZiSzrxBlPMycybLMQVIb7gSYkSzN-m2ITRHwREas12Z6T4Hc3uA44TiM8-McIp2xtcNnN7m76vUO2Tq8pQIyLI7Yym7yZY4Qis_zErrUvhh_bRQ
openUrl ctx_ver=Z39.88-2004&ctx_enc=info%3Aofi%2Fenc%3AUTF-8&rfr_id=info%3Asid%2Fsummon.serialssolutions.com&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Normal+gastric+tissue+Helicobacter+pylori+infection+is+associated+with+epigenetic+age+acceleration%2C+increased+mitotic+tick+rate%2C+tissue+cell+composition%2C+and+Natural+Killer+cell+methylation+alterations&rft.jtitle=bioRxiv&rft.au=Vlasac%2C+Irma+M&rft.au=Christensen%2C+Brock+C&rft.au=Salas%2C+Lucas+A&rft.date=2023-06-29&rft.issn=2692-8205&rft.eissn=2692-8205&rft_id=info:doi/10.1101%2F2023.06.28.546926&rft.externalDBID=NO_FULL_TEXT
thumbnail_l http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/lc.gif&issn=2692-8205&client=summon
thumbnail_m http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/mc.gif&issn=2692-8205&client=summon
thumbnail_s http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/sc.gif&issn=2692-8205&client=summon